Purpose: To compare the oxygen-uptake (${\dot{\mathrm{V}}\mathrm{O}}_2$) kinetics during skating on a treadmill and skating on a slide board and to discuss potential mechanisms that might control the ${\dot{\mathrm{V}}\mathrm{O}}_2$ kinetics responses during skating. Methods: Breath-by-breath pulmonary ${\dot{\mathrm{V}}\mathrm{O}}_2$ and near-infrared spectroscopy–derived muscle deoxygenated hemoglobin and myoglobin ([HHbMb]) were monitored continuously in 12 well-trained, young, long-track speed skaters. On-transient ${\dot{\mathrm{V}}\mathrm{O}}_2$ and [HHbMb] responses to skating on a treadmill and skating on a slide board at 80% of the estimated gas exchange threshold were fitted as monoexponential function. The signals were time-aligned, and the individual [HHbMb]-to-${\dot{\mathrm{V}}\mathrm{O}}_2$ ratio was calculated as the average value from 20 to 120 s after exercise starts. Results: The time constants for the adjustment of phase II ${\dot{\mathrm{V}}\mathrm{O}}_2$ (τ ${\dot{\mathrm{V}}\mathrm{O}}_2$) and [HHbMb] (τ [HHbMb]) were low and similar between slide board and treadmill skating (18.1 [3.4] vs 18.9 [3.6] for τ ${\dot{\mathrm{V}}\mathrm{O}}_2$ and 12.6 [4.0] vs 12.4 [4.0] s for τ [HHbMb]). The $[\mathrm{HHbMb}]:\dot{\mathrm{V}}{\mathrm{O}}_2$ ratio was not different from 1.0 (P > .05) in both conditions. Conclusions: The fast ${\dot{\mathrm{V}}\mathrm{O}}_2$ kinetics during skating suggest that chronic adaptation to skating might overcome any possible restriction in leg blood flow during low-intensity exercise. The ${\dot{\mathrm{V}}\mathrm{O}}_2$ ratio values also suggest a good matching of O₂ delivery to O₂ utilization in trained speed skaters. The similar τ ${\dot{\mathrm{V}}\mathrm{O}}_2$ and τ [HHbMb] values between slide board and treadmill further reinforce the validity of using a slide board for skating testing and training purposes.